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Genome of the Predatory Volute Melo melo Provides Insights into Adaptive Gene Family Diversification in a Basal Neogastropod Lineage

Created on 02 Nov 2025

Authors

Panda, B., Dey, A., Nath, R., Krishnan, A.

Abstract

Neogastropods represent one of the most diverse and ecologically specialized lineages of marine mollusks, yet many of their early-diverging families remain genomically underexplored. Here, we present the first high-contiguity genome of a volutid species, the tropical predatory snail Melo melo, assembled using PacBio HiFi long reads. The final assembly spans 2.29 Gb with high contiguity (N50: 18.6 Mb) and completeness (BUSCO: 92.4%). Phylogenomic analyses place M. melo as an early-branching neogastropod lineage, supporting the basal position of Volutidae within the order. Comparative genomic analyses reveal expansions of gene families associated with digestion and nutrient metabolism (e.g., metallopeptidases), neurotoxicity (conotoxin-like genes), neurotransmission (ion channels), detoxification (cytochrome P450s and ABC transporters), and innate immunity (C1q and LRR-containing proteins), among others. These expansions are consistent with the predatory behavior of M. melo and its adaptation to sediment-rich benthic habitats. Notably, M. melo lacks nacre-related genes but retains several biomineralization-related gene families previously implicated in shell and pearl formation, offering a genomic basis for investigating its distinctive non-nacreous phenotype. Orthology and CAFE analyses further reveal both conserved and lineage-specific dynamics of gene families across the analyzed gastropods. This study fills a key phylogenetic gap in molluscan genomics and establishes Melo melo as a valuable model for understanding the genomic basis of ecological adaptation and trait evolution in basal neogastropods.

Preprint server: bioRxiv
The authors list and abstract were imported from bioRxiv on 02 Nov 2025.

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