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TGF-β and IL-2 differentially shape T follicular regulatory cell differentiation and stability in vitro.

Created on 25 Jun 2026

Authors

Luisa Bach, Yinshui Chang, Olin Arteaga Transito, Mohammadamin Ghasemi, Lisa Maria Steinheuer, Teresa Steffen, Elena De Domenico, Thomas Ulas, F Thomas Wunderlich, Marc D Beyer, Kevin Thurley, Dirk Baumjohann

Published in

Cellular & molecular immunology. Jun 25, 2026. Epub Jun 25, 2026.

Abstract

T follicular helper (Tfh) cells and T follicular regulatory (Tfr) cells play critical roles in regulating the activity of the germinal center (GC), which is essential for the generation of high-affinity antibodies. In the GC, Tfh cells help B cells to proliferate and to differentiate into memory B cells and long-lived plasma cells. In contrast, Tfr cells, a specialized subset of regulatory T cells (Tregs), modulate the humoral immune response by suppressing excessive or autoreactive B-cell activity. Here, we established an in vitro differentiation protocol for mouse CD4⁺ T cells that yielded CXCR5⁺FoxP3⁺ Tfr cells that exhibited a Bcl6hiPD-1hiCD25loGITRint phenotype and were distinct from Treg and Tfh cells. Functionally, in vitro-generated Tfr cells potently suppressed Tfh cell-driven B-cell class switching to IgG1 and downregulated the expression of B-cell costimulatory ligands. While in vitro-generated Bcl6-deficient Tfh cells were impaired in providing help to B cells for efficient class switching to IgG1, in vitro-generated Bcl6-deficient Tfr cells failed to inhibit Tfh cell-driven B-cell class switching to IgG1. Mechanistically, we showed that Tfr cells emerged from FoxP3+ precursors in low-IL-2 environments through a TGF-β- and c-Maf-dependent pathway, allowing for reprogramming and reinforcement of the follicular regulatory cell program in CD4+ T cells in vitro.

PMID:
42342921
Bibliographic data and abstract were imported from PubMed on 25 Jun 2026.

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