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Cell-fate Decisions Mediated by Bistability and Oscillations in the p53-AKT-Mdm2 Network.

Created on 16 Jul 2026

Authors

Yunfeng Zhang, Conghua Wang

Published in

IEEE transactions on computational biology and bioinformatics. Volume PP. Jul 15, 2026. Epub Jul 15, 2026.

Abstract

The tumor suppressor p53 plays a central role in stress-response signaling and exhibits crosstalk with the AKT pathway through the AKT-Mdm2 axis. Although this coupling has been linked to cell-fate choices between survival and death, the dynamical basis of these transitions remains unclear. In this paper, we develop a delay-differential equation model of the p53 AKT-Mdm2 network to study how interlocked feedback loops, together with transcriptional-translational delay, shape p53 dy namics. Bifurcation analysis reveals three dynamical regimes that can be linked to distinct cellular outcomes: a stable low p53 equilibrium (pro-survival), sustained p53 oscillations (repair associated dynamics), and a stable high-p53 equilibrium (pro death). We show that the positive feedback generated by p53 dependent inhibition of AKT and AKT-enhanced Mdm2 activity produces bistability within a certain range of total AKT levels. We further find that increasing the delay destabilizes the low-p53 equilibrium and induces oscillations through a Hopf bifurcation. For the baseline parameter setting, a further increase in delay suppresses the oscillatory state and drives the system toward the high-p53 attractor. These results indicate how AKT-dependent bistability and delay-controlled instability jointly organize state transitions in the p53-AKT-Mdm2 circuit.

PMID:
42455719
Bibliographic data and abstract were imported from PubMed on 16 Jul 2026.

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